Nils Olav Handegard : Curriculum Vitae

Article Preview – The Journal of Ocean Technology. (n.d.). Retrieved August 22, 2022, from https://www.thejot.net/article-preview/
Pedersen, G., Johnsen, E., Khodabandeloo, B., & Handegard, N. O. (2024). Broadband backscattering by Atlantic herring (Clupea harengus L.) differs when measured from a research vessel vs. a silent uncrewed surface vehicle. ICES Journal of Marine Science, fsae048. https://doi.org/10.1093/icesjms/fsae048
Andersen, L. N., Chu, D., Handegard, N. O., Heimvoll, H., Korneliussen, R., Macaulay, G. J., Ona, E., Patel, R., & Pedersen, G. (2023). Quantitative processing of broadband data as implemented in a scientific split-beam echosounder. Methods in Ecology and Evolution, n/a(n/a). https://doi.org/10.1111/2041-210X.14261
Choi, C., Kampffmeyer, M., Handegard, N. O., Salberg, A.-B., & Jenssen, R. (2023). Deep Semisupervised Semantic Segmentation in Multifrequency Echosounder Data. IEEE Journal of Oceanic Engineering, 1–17. https://doi.org/10.1109/JOE.2022.3226214
McQueen, K., Skjæraasen, J. E., Nyqvist, D., Olsen, E. M., Karlsen, Ø., Meager, J. J., Kvadsheim, P. H., Handegard, N. O., Forland, T. N., de Jong, K., & Sivle, L. D. (2023). Behavioural responses of wild, spawning Atlantic cod (Gadus morhua L.) to seismic airgun exposure. ICES Journal of Marine Science, fsad032. https://doi.org/10.1093/icesjms/fsad032
Munnelly, R. T., Castillo, J. C., Handegard, N. O., Kimball, M. E., Boswell, K. M., & Rieucau, G. (2023). Applications and analytical approaches using imaging sonar for quantifying behavioural interactions among aquatic organisms and their environment. ICES Journal of Marine Science, fsad182. https://doi.org/10.1093/icesjms/fsad182
Pala, A., Oleynik, A., Utseth, I., & Handegard, N. O. (2023). Addressing class imbalance in deep learning for acoustic target classification. ICES Journal of Marine Science, fsad165. https://doi.org/10.1093/icesjms/fsad165
Rubbens, P., Brodie, S., Cordier, T., Destro Barcellos, D., Devos, P., Fernandes-Salvador, J. A., Fincham, J. I., Gomes, A., Handegard, N. O., Howell, K., Jamet, C., Kartveit, K. H., Moustahfid, H., Parcerisas, C., Politikos, D., Sauzède, R., Sokolova, M., Uusitalo, L., Van den Bulcke, L., … Irisson, J.-O. (2023). Machine learning in marine ecology: an overview of techniques and applications. ICES Journal of Marine Science, fsad100. https://doi.org/10.1093/icesjms/fsad100
Seri, S. G., Schinault, M. E., Penna, S. M., Zhu, C., Sivle, L. D., de Jong, K., Handegard, N. O., & Ratilal, P. (2023). Characterizing coastal cod vocalization using a towed hydrophone array. ICES Journal of Marine Science, fsad104. https://doi.org/10.1093/icesjms/fsad104
Tenningen, M., Rosen, S., Taraneh Westergerling, E. H., & Handegard, N. O. (2023). How to obtain clear images from in-trawl cameras near the seabed? A case study from the Barents Sea demersal fishing grounds. Fisheries Research, 268, 106856. https://doi.org/10.1016/j.fishres.2023.106856
Van Engeland, T., Bagøien, E., Wold, A., Cannaby, H. A., Majaneva, S., Vader, A., Rønning, J., Handegard, N. O., Dalpadado, P., & Ingvaldsen, R. B. (2023). Diversity and seasonal development of large zooplankton along physical gradients in the Arctic Barents Sea. Progress in Oceanography, 216, 103065. https://doi.org/10.1016/j.pocean.2023.103065
McQueen, K., Meager, J. J., Nyqvist, D., Skjæraasen, J. E., Olsen, E. M., Karlsen, Ø., Kvadsheim, P. H., Handegard, N. O., Forland, T. N., & Sivle, L. D. (2022). Spawning Atlantic cod (Gadus morhua L.) exposed to noise from seismic airguns do not abandon their spawning site. ICES Journal of Marine Science, 79(10), 2697–2708. https://doi.org/10.1093/icesjms/fsac203
Ordoñez, A., Utseth, I., Brautaset, O., Korneliussen, R., & Handegard, N. O. (2022). Evaluation of echosounder data preparation strategies for modern machine learning models. Fisheries Research, 254, 106411. https://doi.org/10.1016/j.fishres.2022.106411
Allken, V., Rosen, S., Handegard, N. O., & Malde, K. (2021). A deep learning-based method to identify and count pelagic and mesopelagic fishes from trawl camera images. ICES Journal of Marine Science, fsab227. https://doi.org/10.1093/icesjms/fsab227
Allken, V., Rosen, S., Handegard, N. O., & Malde, K. (2021). A real-world dataset and data simulation algorithm for automated fish species identification. Geoscience Data Journal, 8(2), 199–209. https://doi.org/10.1002/gdj3.114
Choi, C., Kampffmeyer, M., Handegard, N. O., Salberg, A.-B., Brautaset, O., Eikvil, L., & Jenssen, R. (2021). Semi-supervised target classification in multi-frequency echosounder data. ICES Journal of Marine Science, fsab140. https://doi.org/10.1093/icesjms/fsab140
Handegard, N. O., Algroy, T., Eikvil, L., Hammersland, H., Tenningen, M., & Ona, E. (2021). Smart Fisheries in Norway: Partnership between Science, Technology, and the Fishing Sector. Journal of Ocean Technology, 16(2). https://imr.brage.unit.no/imr-xmlui/handle/11250/2767785
Handegard, N. O., Eikvil, L., Jenssen, R., Kampffmeyer, M., Salberg, A.-B., & Malde, K. (2021). Machine Learning + Marine Science: Critical Role of Partnerships in Norway. The Journal of Ocean Technology, 16(3). https://www.thejot.net/current-issue/
Rodriguez-Pinto, I. I., Rieucau, G., Handegard, N. O., Kimball, M. E., & Boswell, K. M. (2021). Anthropogenic Marsh Impoundments Alter Collective Tendency in Schooling Fish. Estuaries and Coasts. https://doi.org/10.1007/s12237-021-00961-y
Vabø, R., Moen, E., Smoliński, S., Husebø, Å., Handegard, N. O., & Malde, K. (2021). Automatic interpretation of salmon scales using deep learning. Ecological Informatics, 101322. https://doi.org/10.1016/j.ecoinf.2021.101322
Brautaset, O., Waldeland, A. U., Johnsen, E., Malde, K., Eikvil, L., Salberg, A.-B., & Handegard, N. O. (2020). Acoustic classification in multifrequency echosounder data using deep convolutional neural networks. ICES Journal of Marine Science, 77(4), 1391–1400. https://doi.org/10.1093/icesjms/fsz235
Holmin, A. J., Mousing, E. A., Hjøllo, S. S., Skogen, M. D., Huse, G., & Handegard, N. O. (2020). Evaluating acoustic-trawl survey strategies using an end-to-end ecosystem model. ICES Journal of Marine Science. https://doi.org/10.1093/icesjms/fsaa120
Myers, S. C., Thorsen, A., Smoliński, S., Godiksen, J. A., Malde, K., & Handegard, N. O. (2020). An efficient protocol and data set for automated otolith image analysis. Geoscience Data Journal, 7(1), 80–88. https://doi.org/10.1002/gdj3.86
Rodriguez-Pinto, I. I., Rieucau, G., Handegard, N. O., & Boswell, K. M. (2020). Environmental context elicits behavioural modification of collective state in schooling fish. Animal Behaviour, 165, 107–116. https://doi.org/10.1016/j.anbehav.2020.05.002
Allken, V., Handegard, N. O., Rosen, S., Schreyeck, T., Mahiout, T., & Malde, K. (2019). Fish species identification using a convolutional neural network trained on synthetic data. ICES Journal of Marine Science, 76(1), 342–349. https://doi.org/10.1093/icesjms/fsy147
Anders, N., Howarth, K., Totland, B., Handegard, N. O., Tenningen, M., & Breen, M. (2019). Effects on individual level behaviour in mackerel (Scomber scombrus) of sub-lethal capture related stressors: Crowding and hypoxia. PLOS ONE, 14(3), e0213709. https://doi.org/10.1371/journal.pone.0213709
Fields, D. M., Handegard, N. O., Dalen, J., Eichner, C., Malde, K., Karlsen, Ø., Skiftesvik, A. B., Durif, C. M. F., & Browman, H. I. (2019). Airgun blasts used in marine seismic surveys have limited effects on mortality, and no sublethal effects on behaviour or gene expression, in the copepod Calanus finmarchicus. ICES Journal of Marine Science, 76(7), 2033–2044. https://doi.org/10.1093/icesjms/fsz126
Garcia, H. A., Zhu, C., Schinault, M. E., Kaplan, A. I., Handegard, N. O., Godø, O. R., Ahonen, H., Makris, N. C., Wang, D., Huang, W., & Ratilal, P. (2019). Temporal–spatial, spectral, and source level distributions of fin whale vocalizations in the Norwegian Sea observed with a coherent hydrophone array. ICES Journal of Marine Science, 76(1), 268–283. https://doi.org/10.1093/icesjms/fsy127
Johnsen, E., Totland, A., Skålevik, Å., Holmin, A. J., Dingsør, G. E., Fuglebakk, E., & Handegard, N. O. (2019). StoX: An open source software for marine survey analyses. Methods in Ecology and Evolution, 10(9), 1523–1528. https://doi.org/10.1111/2041-210X.13250
Malde, K., Handegard, N. O., Eikvil, L., & Salberg, A.-B. (2019). Machine intelligence and the data-driven future of marine science. ICES Journal of Marine Science. https://doi.org/10.1093/icesjms/fsz057
Seri, S. G., Zhu, C., Schinault, M., Garcia, H., Handegard, N. O., & Ratilal, P. (2019). Long Range Passive Ocean Acoustic Waveguide Remote Sensing (POAWRS) of Seismo-acoustic Airgun Signals Received on a Coherent Hydrophone Array. OCEANS 2019 MTS/IEEE SEATTLE, 1–8. https://doi.org/10.23919/OCEANS40490.2019.8962739
Snowden, D., Tsontos, V. M., Handegard, N. O., Zarate, M., O’ Brien, K., Casey, K. S., Smith, N., Sagen, H., Bailey, K., Lewis, M. N., & Arms, S. C. (2019). Data Interoperability Between Elements of the Global Ocean Observing System. Frontiers in Marine Science, 6. https://doi.org/10.3389/fmars.2019.00442
Stierhoff, K. L., Zwolinski, J. P., & Demer, D. (2019). Distribution, Biomass, and Demography of Coastal Pelagic Fishes in the California Current Ecosystem During Summer 2018 Based on Acoustic-Trawl Sampling. U.S. Dep. Commer, NOAA Tech. Memo, NMFS-SWFSC-613, 81 pp.
Tenningen, M., Pobitzer, A., Handegard, N. O., & de Jong, K. (2019). Estimating purse seine volume during capture: implications for fish densities and survival of released unwanted catches. ICES Journal of Marine Science, 76(7), 2481–2488. https://doi.org/10.1093/icesjms/fsz119
Moen, E., Handegard, N. O., Allken, V., Albert, O. T., Harbitz, A., & Malde, K. (2018). Automatic interpretation of otoliths using deep learning. PLOS ONE, 13(12), e0204713. https://doi.org/10.1371/journal.pone.0204713
Proud, R., Handegard, N. O., Kloser, R. J., Cox, M. J., & Brierley, A. S. (2018). From siphonophores to deep scattering layers: uncertainty ranges for the estimation of global mesopelagic fish biomass. ICES Journal of Marine Science. https://doi.org/10.1093/icesjms/fsy037
Vatnehol, S., & Handegard, N. O. (2018). Echo integration using non-vertical sonar beams: The bias caused by non-uniform distribution of fish within the echo beam. The Journal of the Acoustical Society of America, 144(4), 2160–2165. https://doi.org/10.1121/1.5057120
Vatnehol, S., Peña, H., & Handegard, N. O. (2018). A method to automatically detect fish aggregations using horizontally scanning sonar. ICES Journal of Marine Science, 75(5), 1803–1812. https://doi.org/10.1093/icesjms/fsy029
Zhu, C., Garcia, H., Kaplan, A., Schinault, M., Handegard, N. O., Godø, O. R., Huang, W., & Ratilal, P. (2018). Detection, Localization and Classification of Multiple Mechanized Ocean Vessels over Continental-Shelf Scale Regions with Passive Ocean Acoustic Waveguide Remote Sensing. Remote Sensing, 10(11), 1699. https://doi.org/10.3390/rs10111699
Handegard, N. O., Holmin, A. J., & Rieucau, G. (2017). Method to observe large scale behavioural waves propagating through fish schools using 4D sonar. ICES Journal of Marine Science, 74(3), 804–812. https://doi.org/10.1093/icesjms/fsw211
Handegard, N. O., Tenningen, M., Howarth, K., Anders, N., Rieucau, G., & Breen, M. (2017). Effects on schooling function in mackerel of sub-lethal capture related stressors: Crowding and hypoxia. PLOS ONE, 12(12), e0190259. https://doi.org/10.1371/journal.pone.0190259
Handegard, N. O., Boswell, K., De Robertis, A., Macaulay, G. J., Rieucau, G., & Sivle, L. D. (2016). Investigating the Effect of Tones and Frequency Sweeps on the Collective Behavior of Penned Herring (Clupea harengus). Advances in Experimental Medicine and Biology, 875, 391–398. https://doi.org/10.1007/978-1-4939-2981-8_46
Kruusmaa, M., Rieucau, G., Montoya, J. C. C., Markna, R., & Handegard, N. O. (2016). Collective responses of a large mackerel school depend on the size and speed of a robotic fish but not on tail motion. Bioinspiration & Biomimetics, 11(5), 056020. https://doi.org/10.1088/1748-3190/11/5/056020
Rieucau, G., Holmin, A. J., Castillo, J. C., Couzin, I. D., & Handegard, N. O. (2016). School level structural and dynamic adjustments to risk promote information transfer and collective evasion in herring. Animal Behaviour, 117, 69–78. https://doi.org/10.1016/j.anbehav.2016.05.002
Skern-Mauritzen, M., Ottersen, G., Handegard, N. O., Huse, G., Dingsør, G. E., Stenseth, N. C., & Kjesbu, O. S. (2016). Ecosystem processes are rarely included in tactical fisheries management. Fish and Fisheries, 17(1), 165–175. https://doi.org/10.1111/faf.12111
Trenkel, V. M., Handegard, N. O., & Weber, T. C. (2016). Observing the ocean interior in support of integrated management. ICES Journal of Marine Science: Journal Du Conseil, fsw132. https://doi.org/10.1093/icesjms/fsw132
Ohlberger, J., & Langangen, Ø. (2015). Population resilience to catastrophic mortality events during early life stages. Ecological Applications: A Publication of the Ecological Society of America, 25(5), 1348–1356. https://doi.org/10.1890/14-1534.1
Ressler, P. H., Dalpadado, P., Macaulay, G. J., Handegard, N., & SkernMauritzen, M. (2015). Acoustic surveys of euphausiids and models of baleen whale distribution in the Barents Sea. Marine Ecology Progress Series, 527, 13–29. https://doi.org/10.3354/meps11257
Rieucau, G., Sivle, L. D., & Handegard, N. O. (2015). Herring perform stronger collective evasive reactions when previously exposed to killer whales calls. Behavioral Ecology, arv186. https://doi.org/10.1093/beheco/arv186
Blanchard, A., Hauge, K. H., Andersen, G., Fosså, J. H., Grøsvik, B. E., Handegard, N. O., Kaiser, M., Meier, S., Olsen, E., & Vikebø, F. (2014). Harmful routines? Uncertainty in science and conflicting views on routine petroleum operations in Norway. Marine Policy, 43, 313–320. https://doi.org/10.1016/j.marpol.2013.07.001
Godø, O. R., Handegard, N. O., Browman, H. I., Macaulay, G. J., Kaartvedt, S., Giske, J., Ona, E., Huse, G., & Johnsen, E. (2014). Marine ecosystem acoustics (MEA): quantifying processes in the sea at the spatio-temporal scales on which they occur. ICES Journal of Marine Science: Journal Du Conseil, 71(8), 2357–2369. https://doi.org/10.1093/icesjms/fsu116
Handegard, N. O., Robertis, A. D., Rieucau, G., Boswell, K., & Macaulay, G. J. (2014). The reaction of a captive herring school to playbacks of a noise-reduced and a conventional research vessel. Canadian Journal of Fisheries and Aquatic Sciences, 72(4), 491–499. https://doi.org/10.1139/cjfas-2014-0257
Rieucau, G., Boswell, K. M., De Robertis, A., Macaulay, G. J., & Handegard, N. O. (2014). Experimental Evidence of Threat-Sensitive Collective Avoidance Responses in a Large Wild-Caught Herring School. PLoS ONE, 9(1), e86726. https://doi.org/10.1371/journal.pone.0086726
Rieucau, G., De Robertis, A., Boswell, K. M., & Handegard, N. O. (2014). School density affects the strength of collective avoidance responses in wild-caught Atlantic herring Clupea harengus: a simulated predator encounter experiment. Journal of Fish Biology, 85(5), 1650–1664. https://doi.org/10.1111/jfb.12520
Rieucau, G., Fernö, A., Ioannou, C. C., & Handegard, N. O. (2014). Towards of a firmer explanation of large shoal formation, maintenance and collective reactions in marine fish. Reviews in Fish Biology and Fisheries, 25(1), 21–37. https://doi.org/10.1007/s11160-014-9367-5
Wickham, H. (2014). Tidy Data. Journal of Statistical Software, 59, 1–23. https://doi.org/10.18637/jss.v059.i10
De Robertis, A., & Handegard, N. O. (2013). Fish avoidance of research vessels and the efficacy of noise-reduced vessels: a review. ICES Journal of Marine Science, 70(1), 34–45. https://doi.org/10.1093/icesjms/fss155
Handegard, N. O., Buisson, L. du, Brehmer, P., Chalmers, S. J., De Robertis, A., Huse, G., Kloser, R., Macaulay, G., Maury, O., Ressler, P. H., Stenseth, N. C., & Godø, O. R. (2013). Towards an acoustic-based coupled observation and modelling system for monitoring and predicting ecosystem dynamics of the open ocean. Fish and Fisheries, 14(4), 605–615. https://doi.org/10.1111/j.1467-2979.2012.00480.x
Handegard, N. O., Tronstad, T. V., & Hovem, J. M. (2013). Evaluating the effect of seismic surveys on fish — the efficacy of different exposure metrics to explain disturbance. Canadian Journal of Fisheries and Aquatic Sciences, 70(9), 1271–1277. https://doi.org/10.1139/cjfas-2012-0465
Peña, H., Handegard, N. O., & Ona, E. (2013). Feeding herring schools do not react to seismic air gun surveys. ICES Journal of Marine Science: Journal Du Conseil, 70(6), 1174–1180. https://doi.org/10.1093/icesjms/fst079
Bazazi, S., Pfennig, K., Handegard, N., & Couzin, I. (2012). Vortex formation and foraging in polyphenic spadefoot toad tadpoles. Behavioral Ecology and Sociobiology, 66(6), 879–889. https://doi.org/10.1007/s00265-012-1336-1
Doksaeter, L., Handegard, N. O., Godo, O. R., Kvadsheim, P. H., & Nordlund, N. (2012). Behavior of captive herring exposed to naval sonar transmissions (1.0–1.6 kHz) throughout a yearly cycle. The Journal of the Acoustical Society of America, 131(2), 1632–1642. https://doi.org/10.1121/1.3675944
Doksæter, L., Kvadsheim, P. H., Handegard, N. O., Nordlund, N., & Godø, O. R. (2012). Behavioral Responses of Captive Herring to Sonar Signals (1-1.6 kHz) of a Naval Frigate Throughout a Yearly Cycle. In A. N. Popper & A. Hawkins (Eds.), The Effects of Noise on Aquatic Life (pp. 379–381). Springer New York. https://doi.org/10.1007/978-1-4419-7311-5_86
Handegard, N. O., Boswell, K. M., Ioannou, C. C., Leblanc, S. P., Tjøstheim, D. B., & Couzin, I. D. (2012). The Dynamics of Coordinated Group Hunting and Collective Information Transfer among Schooling Prey. Current Biology, 22(13), 1213–1217. https://doi.org/10.1016/j.cub.2012.04.050
Holmin, A. J., Handegard, N. O., Korneliussen, R. J., & Tjostheim, D. (2012). Simulations of multi-beam sonar echos from schooling individual fish in a quiet environment. The Journal of the Acoustical Society of America, 132(6), 3720–3734. https://doi.org/10.1121/1.4763981
Sivle, L. D., Kvadsheim, P. H., Ainslie, M. A., Solow, A., Handegard, N. O., Nordlund, N., & Lam, F.-P. A. (2012). Impact of naval sonar signals on Atlantic herring (Clupea harengus) during summer feeding. ICES Journal of Marine Science: Journal Du Conseil, 69(6), 1078–1085. https://doi.org/10.1093/icesjms/fss080
Doksaeter, L., Godo, O. R., Handegard, N. O., Kvadsheim, P. H., Lam, F.-P. A., Donovan, C., & Miller, P. J. O. (2009). Behavioral responses of herring (Clupea harengus) to 1–2 and 6–7 kHz sonar signals and killer whale feeding sounds. The Journal of the Acoustical Society of America, 125(1), 554–564. https://doi.org/10.1121/1.3021301
Handegard, N. O., Pedersen, G., & Brix, O. (2009). Estimating tail-beat frequency using split-beam echosounders. ICES Journal of Marine Science, 66, 1252–1258. https://doi.org/doi:10.1093/icesjms/fsp003
Handegard, N. O., & Tjøstheim, D. (2009). The sampling volume of trawl and acoustics: estimating availability probabilities from observations of tracked individual fish. Canadian Journal of Fisheries & Aquatic Sciences, 66, 425–438. https://doi.org/10.1139/F09-004
Pedersen, G., Handegard, N. O., & Ona, E. (2009). Lateral-aspect target-strength measurements of in situ herring (Clupea harengus). ICES Journal of Marine Science, 66, 1191–1196. https://doi.org/10.1093/icesjms/fsp121
Handegard, N. O., & Williams, K. (2008). Automated tracking of fish in trawls using the DIDSON (dual frequency identification sonar). ICES Journal of Marine Science, 65, 636–644. https://doi.org/10.1093/icesjms/fsn029
Hjellvik, V., Handegard, N. O., & Ona, E. (2008). Correcting for vessel avoidance in acoustic abundance estimates for herring using a generalized linear model. ICES Journal of Marine Science, 65. https://doi.org/doi:10.1093/icesjms/fsn082
Handegard, N. O. (2007). Observing individual fish behavior in fish aggregations: Tracking in dense fish aggregations using a split-beam echosounder. The Journal of the Acoustical Society of America, 122(1), 177–187. https://doi.org/10.1121/1.2739421
Ona, E., Godø, O. R., Handegard, N. O., Hjellvik, V., Patel, R., & Pedersen, G. (2007). Silent research vessels are not quiet. The Journal of the Acoustical Society of America, 121, EL145–EL150. https://doi.org/10.1121/1.2710741
Skaret, G., Slotte, A., Handegard, N. O., Axelsen, B. E., & Jørgensen, R. (2006). Pre-spawning herring in a protected area showed only moderate reaction to a surveying vessel. Fisheries Research, 78(2–3), 359–367. https://doi.org/10.1016/j.fishres.2005.11.007
Handegard, N. O., Patel, R., & Hjellvik, V. (2005). Tracking individual fish from a moving platform using a split-beam transducer. The Journal of the Acoustical Society of America, 118(4), 2210–2223. https://doi.org/10.1121/1.2011410
Handegard, N. O., & Tjøstheim, D. (2005). When fish meets a trawling vessel: examining the behaviour of gadoids using a free floating buoy and acoustic split-beam tracking. Canadian Journal of Fisheries and Aquatic Sciences, 62, 2409–2422. https://doi.org/10.1139/F05-131
Jørgensen, R., Handegard, N. O., Gjøsæter, H., & Slotte, A. (2004). Possible vessel avoidance behaviour of capelin in a feeding area and on a spawning ground. Fisheries Research, 69, 251–261. https://doi.org/10.1016/j.fishres.2004.04.012
Patel, R., Handegard, N. O., & Godø, O. R. (2004). Behaviour of herring ( Clupea harengus L.) towards an approaching autonomous underwater vehicle. ICES Journal of Marine Science, 61(7), 1044–1049. https://doi.org/10.1016/j.icesjms.2004.07.002
Handegard, N. O., Michalsen, K., & Tjøstheim, D. (2003). Avoidance behaviour in cod (ıt Gadus morhua) to a bottom-trawling vessel. Aquatic Living Resources, 16, 265–270. https://doi.org/10.1016/S0990-7440(03)00020-2